EPHEMEROPTERA OF THE WORLD

arb.sig. ABC

z   

submarginata Stephens 1835 [Ephemera]

sources:

subjects:

valid names:

invalid synonyms:

Stephens 1835*

Britain

Ephemera submarginata sp.n.

 

Pictet 1843-1845

 

Potamanthus submarginatus comb.n.

 

Walker 1853

Potamanthus submarginatus  

 

Eaton 1871*

 

syn. Leptophlebia helvipes syn.n.

 

McLachlan 1884

Vosges

Leptophlebia submarginata

 

Eaton 1883-1888

Leptophlebia submarginata comb.n.

= Ephemera dispar syn.n.

= Ephemera helvipes syn.n. 

= Potamanthus Geerii syn.n.

= ?Baetis reticulata syn.n.

= Cloeon culiciformis
Walker 1853
(non Linnaeus 1758)

Rostock (1887)1888

Leptophlebia submarginata

= Leptophlebia helvipes

Mocsary 1899

 

 

Klapalek 1906

Chorvatia

?Leptophlebia submarginata

 

Pongracz 1914

 

 

 

Lestage 1917

 

Paraleptophlebia submarginata gen.comb.n.

 

Ulmer 1920:97-144*

-

Paraleptophlebia submarginata

Komarek 1921

Leptophlebia submarginata

 

Ulmer 1929

Paraleptophlebia submarginata

 

Schoenemund 1930

Paraleptophlebia submarginata

 

Allen & Gambles 1932

England

Paraleptophlebia submarginata

 

Eidel 1933

Deutschland

Paraleptophlebia submarginata

 

Mackiewicz-Gutowska 1935*

Poland

Paraleptophlebia submarginata

 

Tiensuu 1935

 

Paraleptophlebia submarginata

 

Mikulski 1936

Paraleptophlebia submarginata

 

Borang 1937

Estonia

Paraleptophlebia submarginata

 

Haberman 1953*

Estonia

Paraleptophlebia submarginata

 

Kimmins 1936:147-151

Britain

Paraleptophlebia submarginata

 

Kimmins 1938:202-209

Britain

Paraleptophlebia submarginata

 

Tiensuu 1939

 

Paraleptophlebia submarginata

 

Kimmins 1939

 

Paraleptophlebia submarginata

 

Kimmins 1942

Paraleptophlebia submarginata

 

Tshernova 1941

n.Russia

Paraleptophlebia submarginata

 

Macan & Macan 1940

r.Avon

Paraleptophlebia submarginata

 

Verrier 1946

ven.variab.

Paraleptophlebia submarginata

 

Macan 1952:363-376*

Paraleptophlebia submarginata

 

Arvy & Gabe 1952

corpora allata

Paraleptophlebia submarginata

 

Zelinka 1953:181-200

Paraleptophlebia submarginata

 

Pleskot 1953

(in detail)

Paraleptophlebia submarginata

 

Pleskot 1953

Wienerwald

Paraleptophlebia submarginata

 

Grandi 1956

genit.

Paraleptophlebia submarginata

 

Jensen C.F. 1956

Denmark

Paraleptophlebia submarginata

Macan 1957:129-156.

Paraleptophlebia submarginata

 

Dudich 1958

Karpaten

Paraleptophlebia submarginata

 

Bogoescu 1958

Paraleptophlebia submarginata

 

Sowa 1959

Paraleptophlebia submarginata

 

Kazlauskas 1959

Lithuania

Paraleptophlebia submarginata

 

Ujhelyi 1959

Paraleptophlebia submarginata

 

Degrange 1960*

reprod.,

Paraleptophlebia submarginata

 

Grandi 1960

Paraleptophlebia submarginata

 

Grandi 1960*

genit.

Paraleptophlebia submarginata

 

Keffermuller 1960

 

Paraleptophlebia submarginata

 

Müller-Liebenau 1960

Eifel

Paraleptophlebia submarginata

 

Ikonomov 1960*

Macedonia

Paraleptophlebia submarginata macedonica

 

Macan 1961:1-64

Britain

Paraleptophlebia submarginata

 

Landa 1962*

Paraleptophlebia submarginata

 

Ikonomov 1964*

Macedonia

Paraleptophlebia submarginata macedonica

 

Gerd & Sokolova 1965

Karelia

Paraleptophlebia submarginata

 

Brekke 1965*

Norvegia

Paraleptophlebia submarginata

 

Langford 1966

Britain

Paraleptophlebia submarginata

 

Ujhelyi 1966

 

Paraleptophlebia submarginata

 

Marchetti & Melone 
& Ramusino 1967

Paraleptophlebia submarginata

 

Illies 1967

 

Paraleptophlebia submarginata

 

Elliott 1967*

Paraleptophlebia submarginata

 

Werner & Werner 1968*

Deutschland

Paraleptophlebia submarginata

 

Landa 1969

Paraleptophlebia submarginata

 

Kimmins 1971

lectotyp.d.

Paraleptophlebia submarginata

 

Kacalova & Skrube 1971*

Latvia

Paraleptophlebia submarginata

 

Jazdewska 1971*

Paraleptophlebia submarginata

 

Lavandier & Dumas 1971:7-23

Paraleptophlebia submarginata

 

Lavandier & Dumas 1971:157-172.

Paraleptophlebia submarginata

 

Sadyrin 1972

Kirovsk.prov.

Paraleptophlebia submarginata

 

Malzacher 1973*

Paraleptophlebia submarginata

 

Müller-Liebenau 1973

DeutschDR

Paraleptophlebia submarginata

 

Dahlby 1973

Norvegia

Paraleptophlebia submarginata

 

Jacob 1974

 

Paraleptophlebia submarginata

= Paraleptophlebia reticulata 

Lavandier & Capblancq 1975

Paraleptophlebia submarginata

 

Sowa 1975

Paraleptophlebia submarginata

 

Brittain 1975

N.Wales

Paraleptophlebia submarginata

 

Zelinka 1976*

Paraleptophlebia submarginata

 

Russev 1977:3-22*

Paraleptophlebia submarginata

 

Kazlauskas 1977

Paraleptophlebia submarginata

 

Hulden & Meinander &
Nyhom & Silfverberg 1977

Finland

Paraleptophlebia submarginata

 

Illies 1978

 

Paraleptophlebia submarginata

= Leptophlebia castanea syn.n.

Alba-Tercedor & Millan 1978

Paraleptophlebia submarginata

 

Belfiore 1978

Italia

Paraleptophlebia submarginata

 

Illies 1978

 

Paraleptophlebia submarginata

 

Koch 1979

Austria

Paraleptophlebia submarginata

 

Ujhelyi 1979

 

Paraleptophlebia submarginata

 

Soldan 1979:22-33

reprod.syst.

Paraleptophlebia submarginata

Soldan 1979*

reprod.sys.

Paraleptophlebia submarginata

 

Soldan 1979:223-230*

spermatozoid

Paraleptophlebia submarginata

 

Belfiore & Giangrande 1979

Paraleptophlebia submarginata

 

Elliott & Humpesch 1980*

oviposition

Paraleptophlebia submarginata

 

Savolainen 1980

Finland

Paraleptophlebia submarginata

 

Puthz 1980

Albania

Paraleptophlebia submarginata

 

Savolainen & Saaristo 1981

Finland

Paraleptophlebia submarginata

 

Alba-Tercedor 1981

España

Paraleptophlebia submarginata

 

Braasch 1981

Iran

Paraleptophlebia submarginata

 

Tanago & Viedma 1981*

Paraleptophlebia submarginata

 

Malzacher 1981:41-72*

s-w.Deutschland

Paraleptophlebia submarginata

 

Sander 1981

Schwarzwald

Paraleptophlebia submarginata

 

Devan 1982*

Czeckoslovakia

Paraleptophlebia submarginata

 

Spuris 1982*

Latvia

Paraleptophlebia submarginata

 

Puig 1983

España

Paraleptophlebia submarginata

 

Thomas & Gadea 1983

France

Paraleptophlebia submarginata

 

Elliott & Humpesch 1983

Paraleptophlebia submarginata

 

Gonzalez del Tanago 1984:IV.Conf.Eph.

Paraleptophlebia submarginata

 

Devan 1984

Slovakia

Paraleptophlebia submarginata

 

Savolainen & Saaristo 1984*

Finland

Paraleptophlebia submarginata

 

Gaino & Belfiore & Spano 1984

Liguria

Paraleptophlebia submarginata

 

Jazdzewska 1984:IV.Conf.Eph.

Paraleptophlebia submarginata

 

Zelinka 1984:IV.Conf.Eph.

Paraleptophlebia submarginata

 

Zurwerra & Tomka 1984

Suisse

Paraleptophlebia submarginata

 

Novikova 1984

n.Urals

Paraleptophlebia submarginata

 

Mol 1984

Netherlands

Paraleptophlebia submarginata

 

Mol 1985

Netherlands

Paraleptophlebia submarginata

 

Landa & Soldan 1985

distrib.

Paraleptophlebia submarginata

 

Sartori & Dethier 1985

Suisse

Paraleptophlebia submarginata

 

Rosillon 1985*

Paraleptophlebia submarginata

 

Jacob 1986*

Paraleptophlebia submarginata

 

Herrera 1986

España

Paraleptophlebia submarginata

 

Devan & Mucina 1986

Paraleptophlebia submarginata

 

Lehrke & Reusch 1987

Saxonia

Paraleptophlebia submarginata

 

Grimm 1987* 

DeutschBR

Paraleptophlebia submarginata

 

Savolainen & Pulkkinen 1987*

Paraleptophlebia submarginata

 

Boev & Bajanov 1987

s.Urals

Paraleptophlebia submarginata

 

Vincon & Thomas 1987*

Paraleptophlebia submarginata

 

Elliott & Humpesch & Macan 1988

Paraleptophlebia submarginata

 

Andrikovicz 1988*

Hungaria

Paraleptophlebia submarginata

 

Landa & Soldan 1989*

Paraleptophlebia submarginata

 

Lewandowski 1989*

Poland

Paraleptophlebia submarginata

 

Bauernfeind 1990*

Austria

Paraleptophlebia submarginata

 

Andrikovics 1991

 

Paraleptophlebia submarginata

 

Andrikovics & Keri 1991

 

Paraleptophlebia submarginata

 

Hefti & Tomka 1991*

Paraleptophlebia submarginata

 

Jazdzewska & Gorczynski 1991:VI.Conf.Eph.

Paraleptophlebia submarginata

 

Preso & Postigo & 
Soto & Luis 1991:VI.Conf.Eph.

Paraleptophlebia submarginata

 

Sartori 1992

Israil

Paraleptophlebia submarginata

 

Studemann & Landolt & 
Sartori & Hefti & Tomka 1992

Paraleptophlebia submarginata

 

Bauernfeind 1992:93-97

Paraleptophlebia submarginata

 

Devan 1993*

Paraleptophlebia submarginata

 

Devan 1993*

Paraleptophlebia submarginata

 

Belfiore 1994*

Italia

Paraleptophlebia submarginata

 

Haybach & Fischer 1994*

Deutschland

Paraleptophlebia submarginata

 

Thomas 1996*

n-e.France

Paraleptophlebia submarginata

 

Parele & Ruzans & 
Spuris 1997*

Latvia

Paraleptophlebia submarginata

 

Moog & al. 1997:VIII.Conf.Eph.

saprob.

Paraleptophlebia submarginata

 

Kluge 1997:233-235

 

Leptophlebia (Paraleptophlebia) submarginata

 

Kluge 1997:176-220

distrib.

Leptophlebia (Paraleptophlebia) submarginata  

 

Juhasz  & Turcsanyi & Kovacs & 
Olajos & Turcsanyi & Kiss 1998

Paraleptophlebia submarginata

 

Godunko 1998*

Ukraina

Paraleptophlebia submarginata

 

Weichselbaumer & Bauernfeind 1999*

Austria

Paraleptophlebia submarginata

 

Kovacs & Ambrus & Bankuti 1999*

Hungaria

Paraleptophlebia submarginata

 

Haybach & Pottgiesser 2000*

N.Westfalen

Paraleptophlebia submarginata

 

Soldan & Puthz 2000*

chromosoms

Paraleptophlebia submarginata

 

Silina & Prokin 2000*

Lipetsk.prov.

Paraleptophlebia submarginata

 

Kazanci 2001:1-80*

Turcia

Paraleptophlebia submarginata

 

Kazanci 2001:1-72 *

Turcia

Paraleptophlebia submarginata

 

Kovacs 2001:163-169*

Hungaria

Paraleptophlebia submarginata

 

Kovacs & Ambrus 2001*

Hungaria

Paraleptophlebia submarginata

 

Kovacs & Juhasz 
& Turcsanyi 2001*

Hungaria

Paraleptophlebia submarginata

 

Kovacs & Ambrus 2002

 

Paraleptophlebia submarginata

 

Kovacs & Ambrus & 
Juhasz 2002:163-167*

Hungaria

Paraleptophlebia submarginata

 

Kovacs & Ambrus & 
Juhasz 2002:169-178*

Hungaria

Paraleptophlebia submarginata

 

Kovacs & Ambrus & Juhasz 2003*

Hungaria

Paraleptophlebia submarginata

 

Kovacs & Bauernfeind 2003*

Hungaria

Paraleptophlebia submarginata

 

Beketov & Kluge 2003

W.Siberia

Leptophlebia (Paraleptophlebia) submarginata

 

Tanatmis 2004*

Turcia

Paraleptophlebia submarginata

 

Tanatmis 2004:45-56*

Turcia

Paraleptophlebia submarginata

 

Ubero-Pascal & Fortuño 
& Puig 2005*

SEM

Paraleptophlebia submarginata

 

Bauernfeind & Kovacs 
& Ambrus 2005*

Hungaria

Paraleptophlebia submarginata

 

Kovacs 2005:101-110*

Hungaria

Paraleptophlebia submarginata

 

Kovacs 2006:143-158*

Hungaria

Paraleptophlebia submarginata

 

Kovacs 2006:325-331*

Hungaria

Paraleptophlebia submarginata

 

Kovacs & Kovacs 2006:159-165*

Hungaria

Paraleptophlebia submarginata

 

Sharapova 2007

W.Siberia

Paraleptophlebia submarginata

 

Giustini & Miccoli & 
De Luca & Cicolani 2008

length/wieght

Paraleptophlebia submarginata

 

Beketov 2008:289-299

Leptophlebia submarginata

 

O'Donnell & Jockusch 2008*

DNA

Paraleptophlebia submarginata

 

Kovacs & Ambrus & Juhasz & Olajos & Szilagyi 2008

Lithuania

Paraleptophlebia submarginata

 

Savolainen 2009:1-35

Finland

Paraleptophlebia submarginata

 

Kovacs 2009:73-85*

Hungaria

Paraleptophlebia submarginata

 

Sroka 2009:XII.Conf.Eph.

mola; food

Paraleptophlebia submarginata

 

Lopez-Rodriguez & Tierno de Figueroa 
& Alba-Tercedor 2010

Paraleptophlebia submarginata

 

Martynov & Godunko 2010

Donetsk.reg.

Paraleptophlebia submarginata

 

Martynov 2011

Kharkov reg.

Paraleptophlebia submarginata

 

Kjaerstad & Webb & Ekrem 2012

N.Norwegia

Paraleptophlebia submarginata

 

Bauernfeind & Soldán 2012

Paraleptophlebia submarginata

 

Martynov 2013(MS)

 

Paraleptophlebia submarginata

 

Cardoni & Tenchini & Ficulle & Piredda & Simeone & Belfiore 2015

DNA

Paraleptophlebia submarginata

 

Martynov 2014:3-18

Donetsk.reg.

Paraleptophlebia submarginata

 

Martynov 2016:86-94.

E.Ukraina

Paraleptophlebia submarginata

 

Bojková & Sroka & Soldán & Namin & Staniczek & Polášek & Hrivniak & Abdoli & Godunko 2018

Iran

Paraleptophlebia submarginata

 

Bauernfeind 2018

Albania

Paraleptophlebia submarginata

 

Righetti 2020:109-122.

Paraleptophlebia submarginata

 

Kluge 2022:151-186.

tarsus

Paraleptophlebia submarginata

 

Le Doaré & Brulin 2023

France

Paraleptophlebia submarginata

 

 

 

Paraleptophlebia submarginata

 

--/  

in Paraleptophlebia/g1

= dispar [Ephemera]

= helvipes [Ephemera]

= reticulata [Baetis]

= geerii [Potamanthus]

= castanea [Leptophlebia]